• Associate Professor, University of South-Eastern Norway, Department of Natural Sciences and Environmental Health, Campus Bø, Norway
• Privatdozent, University of Bonn, Germany
• Arctic-Alpine Ecosystems; Reindeer Research; Alpine Plant Growth
• Biogeography; Dendroecology; Ecoclimatology; Movement Ecology
• Scandies, Greenland, Mediterranean, South Africa
• Co-Chair of German Biogeography Study Group (AK Biogeographie, DGfG)

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Articles in peer-reviewed journals

2024

ALBRECHT, E. C., DOBBERT, S.,  PAPE, R. & LÖFFLER, J. (2024) Complex Environmental Control of Growth in a Dominant Mediterranean-Alpine Shrub Species. Journal of Ecology (accepted)

ALBRECHT, E. C., DOBBERT, S.,  PAPE, R., LÖFFLER, J. (2024) Patterns, Timing, and Environmental Drivers of Growth in two Coexisting Green-Stemmed Mediterranean Alpine Shrubs Species. New Phytologist, 241, 114-130. https://doi.org/10.1111/nph.19285

2023

ALBRECHT, E. C., DOBBERT, S., PAPE, R. & LÖFFLER, J. (2023) Patterns, Timing, and Environmental Drivers of Secondary Growth in two Physiologically Distinct Mediterranean-Alpine Shrub Species. Environmental Research: Ecology, 2, 045002. https://doi.org/10.1088/2752-664X/acffcc

LÖFFLER, J., TSCHORN, K. M., DOBBERT, S., ALBRECHT, E.C., R. PAPE, R. & WUNDRAM, D. (2023) Dataset from Long-Term Air Quality Monitoring in the World Natural Heritage Geirangerfjord, Western Norway (AQM-G). Erdkunde, 77, DP311205. https://doi.org/10.3112/erdkunde.2023.dp.03

ALBRECHT, E. C., DOBBERT, S., PAPE, R., WUNDRAM, D. & LÖFFLER, J. (2023) Annual Ring Width in the Mediterranean-Alpine Shrub Species Cytisus galianoi—Dataset from Long-Term Alpine Ecosystem Research in the Sierra Nevada, Spain (LTAER-ES). Erdkunde; 77, DP311203. https://doi.org/10.3112/erdkunde.2023.dp.01

KÜHNAPFEL, M., ALBRECHT, E. C., DOBBERT, S., PAPE, R., WUNDRAM, D. & LÖFFLER, J. (2023) Annual Ring Width in the Arctic-Alpine Dwarf-Shrub Species Salix herbacea—Dataset from Long-Term Alpine Ecosystem Research in Central Norway (LTAER-NO). Erdkunde; 77, DP311204. https://doi.org/10.3112/erdkunde.2023.dp.02

WEHNER, A., HEIN, N., DOBBERT, S., PAPE, R. & LÖFFLER, J. (2023) Early Snow Melt and Diverging Thermal Constraints Control Body Size in Arctic-Alpine Spiders. Biological Journal of the Linnean Society, 138, 1–13. https://doi.org/10.1093/biolinnean/blac127

2022

DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) On Growth Patterns and Mechanisms in Arctic-Alpine Shrubs. Erdkunde, 76, 199–226. https://doi.org/10.3112/erdkunde.2022.03.04

WEHNER, A., HEIN, N., BECKERS, N., DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) Data from: Early Snow Melt and Diverging Thermal Constraints Control Body Size in Arctic-Alpine Spiders. Dryad Digital Repository. https://doi.org/10.5061/dryad.18931zd1c

DOBBERT, S., ALBRECHT, E. C., PAPE, R. & LÖFFLER, J. (2022) Alpine Shrub Growth Follows Bimodal Patterns Across Biomes—Unexpected Environmental Controls. Communications Biology, 5, 739. https://doi.org/10.1038/s42003-022-03741-x

DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) The Application of Dendrometers to Alpine Dwarf Shrubs—A Case Study to Investigate Stem Growth Responses to Environmental Conditions. Biogeosciences, 19, 1933–1958. https://doi.org/10.5194/bg-19-1933-2022

LÖFFLER, J., ALBRECHT, E. C., DOBBERT, S., PAPE, R. & WUNDRAM, D. (2022) Dendrometer Measurements of Mediterranean-Alpine Dwarf Shrubs and Micro-Environmental Drivers of Plant Growth—Dataset from Long-Term Alpine Ecosystem Research in the Sierra Nevada, Spain (LTAER-ES). Erdkunde, DP311202. https://doi.org/10.3112/erdkunde.2022.dp.01

2021

DOBBERT, S., PAPE, R. & LÖFFLER, J. (2021) How does Spatial Heterogeneity Affect Inter- and Intraspecific Growth Patterns in Tundra Shrubs? Journal of Ecology, 109, 4115–4131. https://doi.org/10.1111/1365-2745.13784

LÖFFLER, J., DOBBERT, S., PAPE, R. & WUNDRAM, D. (2021) Dendrometer Measurements of Arctic-Alpine Dwarf Shrubs and Micro-Environmental Drivers of Plant Growth—Dataset from Long-Term Alpine Ecosystem Research in Central Norway. Erdkunde, DP311201. https://doi.org/10.3112/erdkunde.2021.dp.01

DOBBERT, S., PAPE, R. & LÖFFLER, J. (2021) A New Mechanistic Understanding of Ecophysiological Patterns in a Widespread Alpine Dwarf Shrub—Refining Climate-Growth Relationships. Biogeosciences Discussions 1–40 (Pre-Print).

DOBBERT, S., PAPE, R. & LÖFFLER, J. (2021) Contrasting Growth Response of Evergreen and Deciduous Arctic-Alpine Shrub Species to Climate Variability. Ecosphere, 12 (8), e03688. https://doi.org/10.1002/ecs2.3688 Press Release

DOBBERT, S., PAPE, R. & LÖFFLER, J. (2021) Continent-Wide Modelling of Potential Habitats and Connectivity for the Eurasian Lynx (Lynx lynx) in Europe. Erdkunde, 75, 69–85. https://doi.org/10.3112/erdkunde.2021.02.01

2020

SANDHAGE-HOFMANN, A., LÖFFLER, J., KOTZE, E., WEIJERS, S., WINGATE, V., WUNDRAM, D., WEIHERMÜLLER, L., PAPE, R., DU PREEZ, C. C. & AMELUNG, W. (2020) Woody Encroachment and Related Soil Properties in Different Tenure-Based Management Systems of Semiarid Rangelands. Geoderma, 372, 114399. https://doi.org/10.1016/j.geoderma.2020.114399

LÖFFLER, J. & PAPE, R. (2020) Thermal Niche Predictors of Alpine Plant Species. Ecology, 101, e02891. https://doi.org/10.1002/ecy.2891

2019

HEIN, N., PÉTILLON, J., PAPE, R., FEILHAUER, H., VANSELOW, K. A. & LÖFFLER, J. (2019) Broad-Scale Rather than Fine-Scale Environmental Variation Drives Body Size in a Wandering Predator (Araneae, Lycosidae). Arctic, Antarctic, and Alpine Research, 51, 315–326. https://doi.org/10.1080/15230430.2019.1640039

FRINDTE, K., PAPE, R., WERNER, K., LÖFFLER, J. & KNIEF, C. (2019) Temperature and Soil Moisture Control Microbial Community Composition in an Arctic-Alpine Ecosystem Along Elevational and Micro-Topographic Gradients. ISME Journal, 13, 2031–2043. https://doi.org/10.1038/s41396-019-0409-9

2018

ZIMMERMANN, S., DUBOVYK, O., OLDENBURG, C., PAPE, R. & LÖFFLER, J. (2018) Lichen Cover Mapping in Southern Norway—An Analysis with Remote Sensing and GIS. GIS Science, 2, 60–71.

WEIJERS, S., PAPE, R., LÖFFLER, J. & MYERS-SMITH, I. H. (2018) Contrasting Shrub Species Respond to Early Summer Temperatures Leading to Correspondence of Shrub Growth Patterns. Environmental Research Letters, 13, 034005. https://doi.org/10.1088/1748-9326/aaa5b8

2017

PAPE, R. & LÖFFLER, J. (2017) Determinants of Arctic-Alpine Pasture Resources—The Need for a Spatially and Functionally Fine-Scaled Perspective. Geografiska Annaler: Series A, Physical Geography, 99, 353–370. https://doi.org/10.1080/04353676.2017.1368833

PAPE, R. & LÖFFLER, J. (2016) Towards a Process-Based Biogeography of Reindeer: Scaling Space, Time, and Organizational Levels of Space Use. Norwegian Journal of Geography, 70, 230–246. https://doi.org/10.1080/00291951.2016.1203018

PAPE, R. & LÖFFLER, J. (2016) Broad-Scale Assumptions on Available Pasture Resources and Reindeer´s Habitat Preferences Shown to be Decoupled from Ecological Reality of Arctic-Alpine Landscapes. Erdkunde, 70, 169–192. https://doi.org/10.3112/erdkunde.2016.02.05

PAPE, R. & LÖFFLER, J. (2016) Spatial Patterns of Alpine Phytomass, Primary Productivity, and related Calorific Resources. Ecosphere, 7 (6), e01347. https://doi.org/10.1002/ecs2.1347

2015

PAPE, R. & LÖFFLER, J. (2015) Seasonality of Habitat Selection Shown to Buffer Alpine Reindeer Pastoralism Against Climate Variability. Ecosphere, 6 (12), 260. https://doi.org/10.1890/ES15-00169.1

PAPE, R. & LÖFFLER, J. (2015) Ecological Dynamics in Habitat Selection of Reindeer—An Interplay of Spatial Scale, Time, and Individual Animal´s Choice. Polar Biology, 38, 1891–1903. https://doi.org/10.1007/s00300-015-1750-8

2014

HEIN, N., PAPE, R., FINCH, O.-D. & LÖFFLER, J. (2014) Alpine Activity Patterns of Mitopus morio (Fabricius, 1779) are induced by Variations in Temperature and Humidity at Different Scales in Central Norway. Journal of Mountain Science, 11, 644–655. https://doi.org/10.1007/s11629-013-2913-0

2012

PAPE, R. & LÖFFLER, J. (2012) Climate Change, Land Use Conflicts, Predation and Ecological Degradation as Challenges for Reindeer Husbandry in Northern Europe—What Do We Really Know After Half a Century of Research? Ambio, 41, 421–434. https://doi.org/10.1007/s13280-012-0257-6

2011

LÖFFLER, J., ANSCHLAG, K., BAKER, B., FINCH, O.-D., DIEKKRÜGER, B., WUNDRAM, D., SCHRÖDER, B., PAPE, R. & LUNDBERG, A. (2011) Mountain Ecosystem Response to Global Change. Erdkunde, 65, 191–215. https://doi.org/10.3112/erdkunde.2011.02.06

2010

WUNDRAM, D., PAPE, R. & LÖFFLER, J. (2010) Alpine Soil Temperature Variability at Multiple Scales. Arctic, Antarctic, and Alpine Research, 42, 117–128. https://doi.org/10.1657/1938-4246-42.1.117

2009

PAPE, R., WUNDRAM, D. & LÖFFLER, J. (2009) Modelling Near-Surface Temperature Conditions in High Mountain Environments—An Appraisal. Climate Research, 39, 99–109. https://doi.org/10.3354/cr00795

2008

BÄR, A., PAPE, R., BRÄUNING, A. & LÖFFLER, J. (2008) Growth-Ring Variations of Dwarf Shrubs Reflect Regional Climate Signals in Alpine Environments rather than Micro-Climatic Differences. Journal of Biogeography, 35, 625–636. https://doi.org/10.1111/j.1365-2699.2007.01804.x

FINCH, O.-D., LÖFFLER, J. & PAPE, R. (2008) Assessing the Sensitivity of Melanoplus frigidus (Boheman, 1846) (Orthoptera: Acrididae) to Different Weather Conditions: A Modelling Approach Focussing on Development Times. Insect Science, 15, 167–178. https://doi.org/10.1111/j.1744-7917.2008.00198.x

LÖFFLER, J. & PAPE, R. (2008) Diversity Patterns in Relation to the Environment in Alpine Tundra Ecosystems of Northern Norway. Arctic, Antarctic, and Alpine Research, 40, 373–381. https://doi.org/10.1657/1523-0430(06-097) 

2006

LÖFFLER, J., PAPE, R. & WUNDRAM, D. (2006) The Climatologic Significance of Topography, Altitude and Region in High Mountains—A Survey of Oceanic-Continental Differentiations of the Scandes. Erdkunde, 60, 15–24. https://doi.org/10.3112/erdkunde.2006.01.02

2004

LÖFFLER, J., LUNDBERG, A., RÖSSLER, O., BRÄUNING, A., JUNG, G., PAPE, R. & WUNDRAM, D. (2004) The Central Norwegian Alpine Tree Line under a Changing Climate and Changing Land Use. Norwegian Journal of Geography, 58, 183–193. https://doi.org/10.1080/00291950410002421

LÖFFLER, J. & PAPE, R. (2004) Across Scale Temperature Modelling Using a Simple Approach for the Characterisation of High Mountain Ecosystem Complexity. Erdkunde, 58, 331–348. https://doi.org/10.3112/erdkunde.2004.04.04

PAPE, R. & LÖFFLER, J. (2004) Spatio-Temporal Near-Surface Temperature Variation in High Mountain Landscapes. Ecological Modelling, 178, 483–501. https://doi.org/10.1016/j.ecolmodel.2004.02.019

Other publications

2015

PAPE, R. & LÖFFLER, J. (2015) Mehr als Raum und Zeit—Skalen und die Habitatselektion von Rentieren in Norwegen. Norden, 21, 37–44. Bremen.

2008

LÖFFLER, J. & PAPE, R. (2008) Plant Diversity Patterns and Reindeer Pastoralism in Northern Norwegian Mountain Systems. Colloquium Geographicum, 31, pp. 57–72, Bonn.

2003

LÖFFLER, J., FÜRSTE, C., HEIDER, T., KELCH, T., LÖLKE, K., NAUJOK, J., PAPE, R., SCHADEK, U., SIMON, M. & SPARKE, S. (2003) Aspekte der Landschaftsdegradation auf Island. Oldenburger Geoökologisches Kolloquium, 9, pp. 3–40, Oldenburg.

LÖFFLER, J. & PAPE, R. (2003) Simulation des Energiehaushalts in mittelnorwegischen Hochgebirgslandschaften. Norden, 15, pp. 221–233, Bremen.

2001

LÖFFLER, J., FINCH, O.-D., NAUJOK, J. & PAPE, R. (2001) Möglichkeiten der Integration zoologischer Aspekte in die landschaftsökologische Untersuchung von Hochgebirgen. Methodendiskussion am Beispiel ökologischer Prozesssysteme und Biozönosen. Naturschutz und Landschaftsplanung, 33, 351–357.

LÖFFLER, J., NAUJOK, J., PAPE, R. & RÖSSLER, O. (2001) Ökosysteme und Landschaftsdegradation im Hochgebirge Westgrönlands. Norden, 14, pp. 103–114, Bremen.